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Supports networks that foster communication and new collaborations among scientists, engineers and educators who share a common interest in a new or developing area of science or engineering.

Supports research and research coordination networks focusing on interactions between plants and their symbionts, pathogens and pests and the molecular, genomic, metabolic, cellular, network and organismal processes that mediate these interactions.

Supports research and training on the structure and function of organisms. Core areas supported include development, behavior, neuroscience, physiology, biomechanics and morphology, microbiology, virology and immunology, and plant and animal genomics.

Supports research and training on evolutionary and ecological processes acting at the level of populations, species, communities, ecosystems, macrosystems and biogeographic extents.

Supports planning proposals and major improvements to biological field stations or laboratories in any terrestrial, marine, estuarine or freshwater environment for research and education.

Supports improvements to and digitization of biological collections and collection-based information to advance biology and increase the broader applicability of collections.

Supports research on living systems at the molecular, subcellular and cellular levels. Core areas supported include cellular dynamics and function; genetic mechanisms; molecular biophysics; and systems and synthetic biology.