Division of Integrative Organismal Systems Core Programs (IOS)

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Important Information for Proposers

A revised version of the NSF Proposal & Award Policies & Procedures Guide (PAPPG) (NSF 22-1), is effective for proposals submitted, or due, on or after October 4, 2021. Please be advised that, depending on the specified due date, the guidelines contained in NSF 22-1 may apply to proposals submitted in response to this funding opportunity.

Supports research and training on the structure and function of organisms. Core areas supported include development, behavior, neuroscience, physiology, biomechanics and morphology, microbiology, immunology, virology, and plant and animal genomics.


The Division of Integrative Organismal Systems (IOS) Core Programs support research aimed at understanding why organisms are structured the way they are and function as they do. Proposals are welcomed in all of the core scientific program areas supported by the Division of Integrative Organismal Systems (IOS). Areas of inquiry include, but are not limited to, developmental biology and the evolution of developmental processes, nervous system development, structure, modification, function, and evolution; biomechanics and functional morphology, physiological processes, symbioses and microbial interactions, interactions of organisms with biotic and abiotic environments, plant and animal genomics, and animal behavior. Proposals should focus on organisms as a fundamental unit of biological organization. Principal Investigators are encouraged to apply systems approaches that will lead to conceptual and theoretical insights and predictions about emergent organismal properties. 

Program contacts

Developmental Systems Program Directors
IOSDSC@nsf.gov (703) 292-8417
Behavioral Systems Program Directors
IOSBSC@nsf.gov (703) 292-8423
Phys. & Struct. Systems Program Directors
IOSPSS@nsf.gov (703) 292-8413
Plant Genome Research Program Directors
dbipgr@nsf.gov (703) 292-8420
Neural Systems Program Directors
IOSNSC@nsf.gov (703) 292-8421

Awards made through this program

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Map of recent awards made through this program